Paleobiology

 
 

Paleobiology with Evolutionary Milestones Across Geological Time

Also see:
Tree of Life for Fossil Record
Geological History
 
 

"visit links within the table for more information"


 

 

 

 

 



 


Geologic Time is divided and subdivided into various categories as presented here:
Eons are divided into Eras; Eras are divided Periods; Periods and subperiods are divided into Epochs
(Epochal subdivisions referred to as "ages" are not given in the chart)

 

 

 

 

 

Phanerozoic
EON
(541 mya to present)

"The age of visible life"

ERA
Periods
Epoch
Evolutionary Milestones

Cenozoic Era
(65 mya to today)

Quaternary
(1.8 mya to today)

Holocene
(11 kya to today)

Modern man radiates, "science" appears and eventually computers and the Internet and Iphones become ubiquitous
Plants and animals domesticated at ~ 13 kya.

Pleistocene
(1.8 mya to 11K)

Neandertals appear and disappear; Homo erectus and Homo sapiens appear; Homo sapien speech attained at ~ 75kya. Stone age commences with a human extinction near miss do to continuing ice age.
Neogene
(23 to 1.8 mya

Pliocene
(5 to 1.8 mya)

Ape-like ancestors of modern humans (Hominids), the australopithecines, as well as Homo habilis appear. Land and marine animals including mammals generally modern. Cooler & drier climate reduces tropical plants as deciduous & coniferous forests as well as grasslands & savannas expand.

Miocene
(23 to 5 mya)

Expansive grasslands formed giving rise to new mammal forms, especially grazing horses, rhinoceros, camels, & antelopes, & of course their predators. Many primates & 1st bipedal apes. Mastodons (megafauna) appear late in epoch.
Some 95% of epoch plants are extant today; almost all extant bird families present by epoch end, as well as about half of modern invertebrates.
Marine vertebrates diversify; whales reaching their maximum.
In the seas, vast proliferating kelp and algae fosters new vertebrate and higher mammal (seals) marine life species and associated apex predators like megalodon sharks.
Paleogene
(65 to 23 mya)

Oligocene
(38 to 23 mya)

Evolution of more modern animals continued, particularly among mammals (1st seals) and among angiosperms. Most modern bird forms present. Grazers adapt to apex predators like the saber-toothed cat with longer legs and speed. Oreodonts herebivores abundant.

Eocene
(54 to 37 mya)

Eocene ends with extinction event.
First grasses appear, a resource for herbovores; trees thrive.
Most extant placental mammals appear: modern camels, cats, dogs, horses & rodents & 1st true primates; 1st whales and sea cows.
Earth covered with forests, with rain forests in North America turning to deciduous trees over epoch.
Snakes and turtles abundant and many new avian orders arise.
Insects abundant and appear modern.
Cartilaginous and ray-finned fish thrive worldwide.

Paleocene
(66 to 56 mya)

Flowering plants (angiosperms begin radiation extending through the Eocene, fostered by co-evolution and appearance of bees. Small mammals radiate
Mesozoic Era
(245 to 65 mya)

Cretaceous
(146 to 65 mya

Upper

Ends with Cretaceous - Tertiary (K–T) extinction event. Major extinctions include non avian dinosaurs and ammonites.
Mosasaurs appear late.

Lower
Flowering plants (angiosperms diversify.); lizards; placental animals (early mammals); Also appearing: snakes; social marsupial and primitive placental animals.
New insect forms appear and radiate; pterosaurs common, then decline; Archosaur reptiles and dinosaurs small to huge apex predators common on land, and Chondrichthyes and Actinopterygii fishes in the seas.
Jurassic
(208 to 146 mya)
Upper
Earliest flowering plants (angiosperms) appear at ~ 160 mya or earlier. Appearances include birds; crabs; frogs and salamanders.
Dinosaurs radiate to dominate the land.
 
Middle
Lower
Triassic
(252 to 201 mya)
Upper

Breakup of Pangaea begins
Major extinction event: tabulate corals and conodonts disappear - ammonoids, reptiles and amphibians decimated
Appearances include: dinosaurs; crocodiles; marine reptiles; turtles; Pterosauria
and mammals
Major groups of seed plants appear.

Middle
Lower

Paleozoic Era
(541 to 252 mya)

Permian
(299 to 252 mya)
Lopingian

Permian-Triassic (P-T) extinction event at 251 mya eradicates 95% of all life including graptolites,
blastoids & remaining Proetid trilobites;
all but articulate crinoids dissapear.

Guadalupian
Seed plants are producing large trees.
Cisuralian
~ 10 miilion years for life to recover from P-T extinction.
Carboniferous
(359 to 299 mya)
Pennsylvanian
(323 to 299 mya)

1st Conifers near end of epoch.
Diverse and common amphibians give rise to 1st reptiles (possibly amniotic) at ~ 315 mya - reptiles become dominant tetrapods by end of epoch.
Dense coal forests form, comprising scale trees, ferns, club trees, tree ferns, giant horsetails, cordaites.
Major radiation of winged insects.
Hexapod arthropods large, diverse, and the primary forest herbivores.

Mississippian
(359 to 323 mya)

High diversity of marine life across brachiopods, bryozoans, echinoderms fishes, mollusks, and .
Land plants divide with seed plants to drier areas and lycopods to wetter areas.
Amniotic eggs appear.
Trilobites become scarce.

Devonian
(419 to 359 mya)
Upper
Mass extinction (F-F)
Land colonized by plants and animals. Appearances include: insects; sharks; amphibians (tetrapods); lung fishes and the earliest seed plants. Extensive radiation of fishes.
Rhizodont predatory lobe-finned fishes appear at ~ 377 mya dominate fresh water well into Pennsylvanian.
Middle
Lower
Silurian
(443 to 419 mya)
Pridoli
Jawed fish, cartilaginous fish and vascular plants appear. Primitive terrestrial predators: Arachnids.
Ludlow
Wenlock
Llandovery
Ordovician
(485 to 443 mya)

 

Upper

 

Mass extinction.
First land plants; bryozoans appear. Trilobites begin to specialize.
Stromatolites become rare & mostly replaced by complex reef systems.

 

Middle

 

 

 

Lower

 

Commences with the great Ordovician radiation (Ordovician explosion), as life recovers from the Cambrian – Ordovician extinction event (488 mya).
Cambrian
(541 to 485 mya)
Furongian
Cambrian – Ordovician event ended the Cambrian Period, where many brachiopods and conodonts perished, and trilobites were severely reduced.
Series 3
Appearances include: vertebrates; jawless fish; small shelly animals mostly extinct by end of early Cambrian (End-Botomian mass extinction); conodonts; trilobites radiate repeatedly and reach their peak diversity.
Series 2
 
Terreneuvian
Cambrian Explosion, the 1st major radiation of animals when most phyla appear. Appearance of hard parts and vision - macroscopic fossils become common.

Proterozoic
EON

(2500 to 541 mya)

Neoproterozoic
(1000 to 541 mya)
- Late

Ediacaran or Vendian
(635 to 541 mya)

Extinction at end of Ediacaran.
Appearance of Tommotian mineralized Fauna (small shelly animals) at ~ 550 mya: they radiate worldwide.
Macroscopic, soft-bodied organisms radiating, the oldest metazoan (multicellular animals) - fossils known as the Ediacaran Biota, including trilobitamorphs, poriferans and other enigmatic forms.


Stromatolites diminishing further, possibly providing food for herbivorous eukaryotes.

Cryogenian
(850 to 635 mya)

Tonian
(1000 to 850 mya)

Chloroblasts arise from cyanobacteria through endosymbiosis at ~ 1000 mya.
Acritarchs radiate, becoming widespread & some perhaps eukaryotic and photosynthetic dinoflagellates or eukaryotic protists.
Mesoproterozoic
(1600 to 1000 mya)
- Middle

Stenian
(1200 to 1000 mya)

 

Ectasian
(1400 to 1200 mya)

 

Calymmian
(1600 to 1400 mya)

Free atmospheric oxygen build-up continues desimating some prokaryotic bacteria, but enabling replacement by newly evolved eukaryotic forms, including photosynthetic multicellular algae.
Paleoproterozoic
(2500 to 1600 mya)
- Early

Statherian
(1800 to 1600 mya)

More complex single-celled life with aerobic metabolism begin diversification.
Eukaryotic mitochondria evolve.
Approximate peak of stromatolites with cyanobacteria oxygenating the atmosphere.
Fossilized filamentous algae (eukaryote) at ~ 1700 mya.

Orosirian
(2050 to 1800 mya)

Banded iron formation diminishes allowing atmosphere to oxygenate rapidly, reaching ~ 15% at ~ 1800 mya.
Rhyacian
(2300 to 2050 mya)
The oldest known potential multicellular eukaryote is Grypania spiralis, appearing as a coiled algae in 2100 mya banded iron formations in Michigan.
Earliest known single-celled eukaryote fossils are acritarchs, which become widespread at ~ 2100 mya. acritarchs are most common fossils of late Proterozoic.
Siderian
(2500 to 2300 mya)
Banded iron formation accelerates at ~ 2400 mya, continuing at high rate until diminishing at ~ 1800 mya -- the rusting of the seas.
Archaean
EON

(4000 to 2500 mya)

Neoarchean
(2800 to 2500 mya)

Molecular fossils from Australia suggest eukaryotes appeared at ~ 2700 mya, but this is not a widely accepted view, with other estimates at ~ 3500 mya.
Atmospheric oxygen only ~ 1%.

Mesoarchean
(3200 to 2800 mya)
Prokaryotes dominate (Eubacteria and Archaea); simple cell forms generate extensive stromatolite reef systems. First acritarch microscopic fossils.
1st substantial free oxygen from photosynthetic archaea and bacteria at ~ 3000 to 2300 mya, after which free oxygen produced by these prokaryotes combined with dissolved iron in the oceans to form banded iron formations until ~ 2000 to 1300 mya -- the so-called rusting of the Earth.
Paleoarchean
(3600 to 3200 mya)

Primitive Eukaryotes as early as ~ 3500 mya after endosymbiosis.
Oxygenic photosynthetic cyanobacteria appear ~ 3500 mya.
Oldest fossils - Apex Chert at ~ 3550 mya & Strelly Pool at ~ 3430 mya in Pilbara, Western Australia

Eoarchaean
(4000 to 3600 mya)

First banded iron formation at ~ 3700 mya.
Oxygenic photosynthetic prokaryotic bacteria appear at ~ 3500 mya.

First life appears
, at ~ 3800 mya & is Archaea or Bacteria, chemotrophic, anerobic, asexual, prokaryotes, fairly soon after end of earth bombardment.
Oldest sedimentary rocks ~ 3800 mya.
Earth's crust cooled by ~ 4000 mya, but atmosphere comprised volcanic gases and minimal oxygen. The 1st oceans were formed.

Hadean
EON

(4567 to 4000 mya)
Lower Imbrian
(4100 to 4000 mya)

Self replicating RNA molecules may might have appeared at ~ 4000 mya.
Late heavy bombardment from space ends at ~ 4000 mya ends, setting stage for life to appear.
Earth's oldest surviving rock from Canada dated at ~ 4030 mya.
Nectarian
(4300 to 4100 mya)
Nectarian begins with so-called Late Heavy Bombardment of Earth from space.
Basin Groups
(4500 to 4300 mya)
Name derived from groupings of major impact crators.
Cryptic
(4567 to 4500 mya)
Earth's environment exceedingly hostile to life as we know it.
Meager geological evidence survives from this time, having been destroyed by bombardment of earth from space projectiles, including the one that putatively formed the moon.
Earth forms at ~ 4567 mya.


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