Osteolepis macrolepidotus
Class
Sarcopterygii, Infraclass Tetrapodomorpha, Superorder Osteolepidida,
Order Osteolepiformes, Family
Osteolepidae
Geological
Time: Middle Devonian
Size: 175
mm in length
Fossil Site:
Cruday Quarry, Orkney, Scotland
Description:
Osteolepis is a member of Order Osteolepiformes within Vertebrata
Class Sarcopterygii, the so-called lobe-finned fish. While they
are bony fish, they are formally considered in classical cladistics
to be terrestrial vertebrates and are among the many ancestors
of tetrapods
that conquered the land late in the Devonian Period (~
360 million years ago). Osteolepis is noted for having scales
and plates on its head that were covered with a thin layer of
cosmine scales (a spongy but bony material).
Osteolepiformes
is also included in an unranked (but believed monophyletic)
clade Rhipidistia made up of Tetrapoda and still
extant lungfishes and Coelacanths. Traits that place Osteolepis
within the lobe-finned fish are paired lobed fins and posterior
nasal passages called choanae between the nasal cavity and
the throat in tetrapods, including humans, that allow breathing
when
the mouth is closed. Osteolepis is also noted for think rhombic-shaped
as seem in the fossil images here. These Devonian fish had
many characteristics that would be largely retained when the
first
fish walked ashore.
Images
by permission from Fossil Mall
(CC BY-NC 4.0) Also
see: Sarcopterygii Fish
Fossils Osteolepis panderi
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