The
Aglaspida (Aglaspids) are an unranked (incertae
sedis) clade of early arthropods that due to their resemblance
to horseshoe crabs were once believed to ancestral horseshoe crabs,
and were included with the chelicerata. Most recently, aglaspids
are held to be distinct group, possibly closely related trilobites,
and possibly linking
the trilobites with the Chelicerata. Although aglaspid fossils
are distributed worldwide, they are relatively uncommon in the
fossil record. They are, in fact, one of the largest non-trilobite
arthropod groups in the fossil record.
Aglaspids
had 8 to 12 pairs of appendages and a prominent telson. These
morphological characteristics have caused considerable support
for Aglaspids being the maker of Protichnites ichnofossils. Protichnites
of the Upper Cambrian Mount
Simon Sandstone in Wisconsin have been suggested as the first
footprints on land in the fossil record, possibly marking the
transition to terrestrial life that took tens of millions years
more to complete.