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The
phylogenetically placement of the trilobites within Phylum
Arthropoda remains problemmatic, as indeed are the relationships
among all the early arthropods. Based on some cladistic analyses,
the Helmetiidae and Naraoiidae
are considered by some to be the early arthropods known from
fossils that are phylogenetically the closest to Trilobita;
sometimes a particular Naraoiidae is placed in another group,
the Tegopeltidae. The Naraoiidae is
a family of early arthropods resembling trilobites in appearance.
Because Naraoids had an entirely non-mineralized exoskeletons,
they are often referred to as soft-bodied trilobites. These
Naraoids
are known only from the early and middle Cambrian, and mainly
from the Maotianshan
Shales of Chengjiang in China, the Burgess
Shale of British Columbia, Canada, and, though rare, the House
Range of Utah (including at least the Marjum
and Weeks Formations). Trilobitomorpha
is a term that decreasingly used as a subphylum of Arthropoda
that encompasses trilobita together with other arthropods that
have at least some trilobite characteristics. Originally a variety
of peculiar forms, mostly from the lower Cambrian, were included
as the Class Trilobitoidea. However, the many species that are
known only from fossils do not appear to be closely related
to
the trilobites or often even to each other either, and are now
generally placed in separate subphyla or superclass when classified
at all. The phylogeny sensu Hou & Bergström (1997) below
shows one possible arrangement of various trilobite relatives.
The
phylogeny sensu Hou & Bergström (1997) below
shows one possible arrangement of various trilobite
relatives.
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< LAMELLIPEDIA sensu Hou & Bergstrom, 1997
|-- ? MARRELLOMORPHA
| |-- ? MARRELLIDA
| | `--? Marrellidae
| `-- ? MIMETASTERIDA
| `-- ? Mimetasteridae?
`-- ARTIOPODA
|-- ? NECTOPLEURA
| |-- ? NECTASPIDA
| | |-- ? Orientellidae
| | `--+-- ? Liwiidae
| | `-- ? Naraoiidae
| |-- ? VACHONISIIDA
| | `-- ? Vachonisia rogeri
| |-- ? ACERCOSTRACA
| |-- ? HALICYNA
| `-- ? RETIFACIIDA
| `-- ? Retifaciidae
`--+-- ? CONCILITERGA
| `-- ? HELMETIIDA
| |-- ? Helmetiidae
| `--+-- ? Skioldiidae
| `--+-- ? Tegopeltidae
| `-- ? Saperiidae
|-- ? PETALOPLEURA
| |-- ? XANDARELLIDA
| | |-- ? Xandarellidae
| | `-- ? Almeniidae
| |-- ? CHELONEILLIDA
| | `-- ? Cheloniellidae
| `-- ? SINOBURIIDA
| `-- ? Sinoburiidae
|-- TRILOBITA
`--+-- ? XENOPODA
| |-- ? EMERALDELLIDA
| | `-- ? Emeraldellidae
| `-- ? LIMULAVIDA
| `-- ? Sidneyiidae
|-- ? AGLASPIDA (AGLASPIDIDA)
| |-- ? AGLASPIDIDA
| | |-- ? Aglaspididae
| | `-- ? Lemoneitidae
| `-- ? STRABOPIDA
| `-- ? Strabopidae [Paleomeridae]
`-- CHELICERATA
-
Hou
& Bergström, 1997: Arthropods of the Lower
Cambrian Chengjiang fauna, southwest China. --Fossils
& Strata, Number 45, 22nd December 1997, pp.
1-116
-
Hou,
X. G.; Bergstrom J, (1991) The arthropods of the
Lower Cambrian Chengjiang fauna, with relationship
and evolutionary significance, p. 179-187. In A.
M. Simonetta and S. Conway Morris (eds.), The Early
Evolution of Metazoa and the Significance of Problematic
Taxa. Cambridge University Press, Cambridge.
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